Cover Page

CONTENTS

SECTION I: NUTRITION AND FEEDS FOR HORSES

Chapter 1. WATER, ENERGY, PROTEIN, CARBOHYDRATES, AND FATS FOR HORSES

NUTRIENTS

WATER

DIETARY ENERGY

PROTEIN

CARBOHYDRATES

FATS

Chapter 2. MINERALS FOR HORSES

CALCIUM AND PHOSPHORUS

SODIUM CHLORIDE SALT

POTASSIUM

MAGNESIUM

SULFUR

SELENIUM AND VITAMIN E

IODINE

COPPER

MOLYBDENUM

ZINC

MANGANESE

IRON

COBALT

FLUORIDE

Chapter 3. VITAMINS FOR HORSES

VITAMIN A

VITAMIN D

VITAMIN K

THIAMIN (VITAMIN B1)

RIBOFLAVIN (VITAMIN B2)

NIACIN

PANTOTHENIC ACID

PYRIDOXINE (VITAMIN B6)

BIOTIN

FOLACIN

COBALAMIN (VITAMIN B12)

CHOLINE

VITAMIN C (ASCORBIC ACID)

Chapter 4. HARVESTED FEEDS FOR HORSES

HAYS FOR HORSES

CEREAL GRAINS FOR HORSES

HIGH-MOISTURE FEEDS FOR HORSES

VITAMIN-MINERAL SUPPLEMENTS FOR HORSES

PROTEIN SUPPLEMENTS FOR HORSES

FAT AND OIL SUPPLEMENTS FOR HORSES

GRAIN MIXES AND COMPLETE FEEDS FOR HORSES

BY-PRODUCT FEEDS FOR HORSES

FEED ADDITIVES FOR HORSES

Chapter 5. PASTURE FOR HORSES

HORSE PASTURE PURPOSE AND BENEFIT

HORSE GRAZING BEHAVIOR

GRAZING MANAGEMENT

PASTURE FERTILIZATION

WEED CONTROL

PLANTING PASTURES

Chapter 6. DIET EVALUATION, FORMULATION, AND PREPARATION FOR HORSES

WHY DIET EVALUATION AND FORMULATION ARE NEEDED

INFORMATION NEEDED FOR DIET EVALUATION AND FORMULATION

DIET EVALUATION AND FORMULATION PROCEDURES

PREPARATION OF GRAIN MIXES

Chapter 7. MINIMIZING HORSE FEEDING COSTS

BUYING FEED ECONOMICALLY

DETERMINING LEAST-COST FEED

MINIMIZING WASTE

SECTION II: FEEDING AND CARE OF HORSES

Chapter 8. GENERAL HORSE FEEDING PRACTICES

WATER FEEDING

FORAGE FEEDING

GRAIN FEEDING

FEEDING FREQUENCY

MONITORING HORSES’ NUTRITIONAL AND HEALTH STATUS

GROUP SOCIALIZATION, COMMUNICATION, AND FEEDING

CHANGING DIETS

Chapter 9. GENERAL HORSE CARE MANAGEMENT PRACTICES

INTERNAL PARASITE PROBLEMS

INTERNAL PARASITE CONTROL

EXTERNAL-PARASITE PROBLEMS AND CONTROL

INFECTIOUS-DISEASE PROBLEMS AND VACCINATION PROGRAMS

DENTAL PROBLEMS AND CARE

FOOT CARE

SKIN AND HAIR-COAT CARE

RECORD KEEPING

HOUSING FOR HORSES

FENCING FOR HORSES

SUPPLEMENTAL READING RECOMMENDATIONS

Chapter 10. FEEDING IDLE AND WORKING HORSES

FEEDING HORSES FOR MAINTENANCE OR WORK

COLD WEATHER CARE OF HORSES

HOT WEATHER CARE OF HORSES

AGED HORSES

Chapter 11. FEEDING AND CARE OF HORSES FOR ATHLETIC PERFORMANCE

ENERGY PRODUCTION AND UTILIZATION FOR ATHLETIC PERFORMANCE

DIETS AND SUPPLEMENTS FOR ATHLETIC PERFORMANCE

FEEDING AND SUPPLEMENTS BEFORE ATHLETIC PERFORMANCE

WATER AND ELECTROLYTES FOR ATHLETIC PERFORMANCE

TRAINING TO INCREASE FITNESS FOR ATHLETIC PERFORMANCE

FITNESS EVALUATION FOR ATHLETIC PERFORMANCE

DISEASES DUE TO ATHLETIC PERFORMANCE

INJURIES DUE TO ATHLETIC PERFORMANCE

MANAGEMENT SUMMARY FOR ATHLETIC PERFORMANCE

SELECTING THE EQUINE ATHLETE

SUPPLEMENTAL READING RECOMMENDATIONS

Chapter 12. BREEDING STALLION FEEDING AND CARE

FEEDING BREEDING STALLIONS

EXERCISE FOR BREEDING STALLIONS

BREEDING EVALUATION OF STALLIONS

BREEDING STALLION USAGE

CASTRATION

MASTURBATION

SUPPLEMENTAL READING RECOMMENDATIONS

Chapter 13. BROODMARE FEEDING AND CARE

VACCINATION AND INTERNAL-PARASITE CONTROL PROGRAMS FOR BROODMARES

FEEDING BROODMARES

NUTRITION EFFECTS ON MARE REPRODUCTION

NUTRITION EFFECTS ON MARE’S MILK PRODUCTION AND COMPOSITION

NUTRITION PROBLEMS IN BROODMARES

MARE’S ESTROUS CYCLE

ESTROUS CYCLE INDUCEMENT OF MARES

PREBREEDING EXAMINATION OF MARES

BREEDING MARES

FOAL-HEAT BREEDING

DIAGNOSING PREGNANCY AND FETAL VIABILITY IN MARES

ABORTION INDUCEMENT IN MARES

TWINNING IN MARES

SUPPLEMENTAL READING RECOMMENDATIONS

Chapter 14. MARE AND FOAL CARE AT FOALING

GESTATION LENGTH IN MARES

PREPARATIONS FOR FOALING

PREDICTING FOALING TIME

INDUCING FOALING

FOALING

BEHAVIOR OF THE MARE AND FOAL AFTER FOALING

NEONATAL FOAL CARE

MARE CARE AFTER FOALING

MARE INJURIES IN FOALING

SUPPLEMENTAL READING RECOMMENDATIONS

Chapter 15. GROWING HORSE FEEDING AND CARE

FEEDING GROWING HORSES

WEANING

GROWTH PROMOTANTS FOR HORSES

EXERCISE FOR GROWING HORSES

VACCINATION AND INTERNAL-PARASITE CONTROL FOR GROWING HORSES

ORPHAN AND EARLY-WEANED FOAL FEEDING AND CARE

Chapter 16. DEVELOPMENTAL ORTHOPEDIC DISEASES IN HORSES

EFFECTS OF DEVELOPMENTAL ORTHOPEDIC DISEASES

CAUSES OF DEVELOPMENTAL ORTHOPEDIC DISEASES

NONSURGICAL MANAGEMENT OF DOD

SUPPLEMENTAL READING RECOMMENDATIONS

Chapter 17. FEEDING AND CARE OF HORSES WITH HEALTH PROBLEMS

INADEQUATE FEED CONSUMPTION

HOOF DEFECTS

HEAVES (CHRONIC OBSTRUCTIVE PULMONARY DISEASE)

DIARRHEA

INTESTINAL IMPACTION

INTESTINAL CALCULI (STONES)

INTESTINAL REMOVAL OR DYSFUNCTION

RECTAL/VAGINAL SURGERY OR LACERATIONS

LIVER DISEASE

KIDNEY FAILURE

URINARY TRACT CALCULI (BLADDER STONES)

HEART DISEASE

POTASSIUM INDUCED PERIODIC PARALYSIS

Chapter 18. PLANT POISONING OF HORSES

SALIVATION-INDUCING PLANTS

COLIC- AND DIARRHEA-INDUCING PLANTS

PRIMARY PHOTODERMATITIS-INDUCING PLANTS

LIVER DISEASE-INDUCING PLANTS

NEUROLOGIC DISEASE-INDUCING PLANTS

LAMENESS- AND MUSCLE WEAKNESS-INDUCING PLANTS

TREATMENT OF CHRONIC SELENIUM EXCESS

ANEMIA-INDUCING PLANTS

TERATOGENIC PLANTS

SUDDEN DEATH-INDUCING PLANTS

SUPPLEMENTAL READING RECOMMENDATIONS

Chapter 19. FEED-RELATED POISONINGS OF HORSES

MYCOTOXIN POISONING

IONOPHORE ANTIBIOTIC POISONING IN HORSES

FORAGE POISONING (BOTULISM) IN HORSES

LEAD POISONING IN HORSES

BLISTER BEETLE POISONING IN HORSES

GOSSYPOL POISONING

NITRATE POISONING

Chapter 20. BEHAVIORAL PROBLEMS IN HORSES

ESCAPE VICES

ORAL VICES

FLIGHT OR FIGHT VICES

SUPPLEMENTAL READING RECOMMENDATIONS

GLOSSARY

APPENDIX TABLES:

INDEX

Title Page

DEDICATION

Dedicated to the memory of

Two outstanding horsemen:

Everette Corwin Lewis, my father, whom I didn’t have the pleasure of knowing and Lee M. Brunk, my father-in-law, whom I did.

Two great cattlemen:

Del J. and Dorcas M. Bigelow, my grandparents, who taught me much about life

and

Veterinary editor and friend to the profession and those they serve, Kit (Christian C. Febiger) Spahr, Jr., editor of the first edition of this book.

All of whom live on in the memory of those who knew them and/or have benefited from the many contributions they left us.

PREFACE

There has been an extensive amount of research conducted, as well as experiences and recommendations reported, on all aspects of feeding and caring of the horse. Much of this information is accurate and applicable, however, some is not. Some must be interpreted or combined with other information to be directly applicable and useable. Results with other species of animals are sometimes reported for horses. Often these are found not to be applicable for horses. Some experiences, recommendations, and “old wives’ tales” are well proven not to be true, or to be true only under certain circumstances, but unfortunately are often repeated by those unaware of studies or experiences refuting them. Occasionally, financial gain rather than scientific validity appears to be the motive behind information and recommendations given. Generally, there is some factual basis for the statements or implications made, but what has been demonstrated is not understood, is misrepresented, is exaggerated, or most often, is extrapolated to circumstances to which they either don’t apply or to which there is no evidence that they do. This book is an effort to alleviate these shortcomings in bringing together research results, experiences, and recommendations in as accurate, complete, and as useable a manner as possible for horse owners, who are not veterinarians or nutritionists.

For the veterinarian and nutritionist the referenced textbook EQUINE CLINICAL NUTRITION: Feeding and Care is more useful. It provides the details necessary for a more complete understanding, for diagnosing, and for both dietary as well as medical management necessary to most successfully treat nutritional and feeding-related diseases affecting horses. Dietary management of non-feeding related diseases for both sick horses and foals are also described, including fluid and electrolyte therapy, and oral and intravenous feeding. Studies confirming the validity and basis for information and recommendations given are described and referenced so additional details can be found if wanted. FEEDING AND CARE OF THE HORSE is an abridgment of this more medically oriented textbook. Research details and references are omitted allowing information to be presented more succinctly. This greatly assists in making the information easier to find, understand and use. There is minimal use of medical and other words not common to non-health professionals. However, at times their use is necessary. In addition, there are some words common to some but not others, and that have different meanings to different people and in different situations. Because of this, a glossary of over 750 words has been added to this book.

The book is divided into three sections. In the first section on “Nutrition and Feeds for Horses” emphasis is on the nutrients needed by the horse and their many sources. In the second section on “Feeding and Care of Horses” feeding to meet these needs, as well as additional aspects of care, are described. The last section contains the glossary and appendix tables which provide information referred to throughout this book and needed repeatedly in feeding and caring for horses.

The book begins with a listing of chapter titles and their location. Each chapter begins with a detailed table of contents for quickly finding specific topics of interest and items involved with or described under that topic. Most chapters end with a description of additional information recommended on topics covered in that chapter.

Chapters 1, 2, and 3 describe the sources, utilization, and functions of nutrients by the horse, and causes, effects, and treatment of deficiencies and excesses of these nutrients – water and energy-providing nutrients in Chapter 1, and the many minerals and vitamins needed, harmful, and given to horses in Chapters 2 and 3, respectively. The multitude of different feeds, both harvested and pasture, that provide these nutrients are described in Chapters 4 and 5. No topic in feeding horses is of greater concern than that of the various feeds that may be used, or are harmful to horses. These feeds include types, forms and cuttings of hay; the many different cereal grains; high-moisture harvested feeds; vitamins, minerals, protein and fat supplements; commercially available feed mixtures; and various by-product feeds, and feed additives fed to horses. However, for many horses the major and often only feed needed is pasture forage for which its types, planting, management and benefit are described. How to determine the nutrient content of the horse’s diet and, therefore, if it is providing inadequate or excessive amounts of each nutrient, and how to mix feeds together so that nutrients low in one feed are balanced by feeding another feed higher in these nutrients, and thus prevent or correct nutrient deficiencies and excesses are described in Chapter 6. How to do this as economically as possible is described in Chapter 7.

The use of the information in Section I on “Nutrition and Feeds” to feed and care for the horse is described in Section II. General feeding and management practices for all horses are given in Chapters 8 and 9. This includes needs, methods and frequency of feeding different types of feeds, the description, harm and control of internal and external parasites and infectious diseases, dental, foot and hair coat problems and care, recordkeeping, housing, and fences. Specific feeding and care for each stage of life, activity and environment are given in Chapters 10 through 15. This includes during hot or cold weather, and for idle, working and aged horses, for athletic performance, for breeding stallions, broodmares and growing horses including orphans. Nutrients, training, fitness evaluation, diseases due to and selecting horses for athletic performance are also described in Chapter 11. Chapters 12 and 13 also include breeding evaluation, procedures and problems, including nutritional effects on reproduction and on milk production and composition. Mare and foal care at foaling, including their behavior, and maternal aggression and complications are described in Chapter 14. Following Chapter 15 describing feeding and care of growing horses, the most common problems occurring in fast growing light horse breeds, developmental orthopedic diseases, are described in Chapter 16.

A tremendous thanks to Dr. Tony Knight for the extensive and thorough compilation given in Chapter 18 of all plants growing in North America confirmed to cause poisoning of horses. The plants are presented according to their major detrimental effect on the horse. This presentation greatly assists in knowing which plants, if any, to consider as a cause of abnormalities in horses. Dr. Knight–s extensive training, experience and expertise in internal medicine combined with his life-long avocation — botany, have resulted in an extremely useful source of information never before available. For each plant a description for its identification, along with colored pictures, its toxic principle and its effects, clinical signs, diagnosis, treatment, and means of preventing poisoning of horses are given. Similar information is given in Chapter 19 on feeds which, in contrast to poisonous plants, are intended for horses, but which under certain circumstances may cause poisonings. These include toxins produced by molds including: fescue poisoning, moldy corn disease, and grass staggers. Poisonings due to antibiotics in feeds intended for other species, botulism, lead and blister beetles are also covered. Cottonseed and nitrate poisonings are described because they are often of concern, although they rarely if ever cause poisoning of horses.

Behavioral problems in horses along with their causes, treatment, and prevention are described in Chapter 20. These include a number of escape, oral, and flight or fight vices. Most are caused by feeding and care without consideration of the horse's psychological needs. As described in this chapter, understanding and caring for these needs will prevent or eliminate many of these vices and are as important in maintaining the horse’s health, happiness, performance, and enjoyability as is maintaining the horse’s physiological well-being as described in the previous chapters of this book.

It is intended and hoped that this book provide an accurate and useful compilation and transmission of the multitude of valid and useful information available on feeding and care of the horse to those providing this care.

Topeka, Kansas

LON D. LEWIS

Section One

NUTRITION AND FEEDS FOR HORSES

Chapter 1

WATER, ENERGY, PROTEIN, CARBOHYDRATES, AND FATS FOR HORSES

Nutrients
Water
Needs
Deficiency
Quality
Dietary Energy
Sources and Use
Needs
Deficiency
Excess
Protein
Needs
Nonprotein Nitrogen (Urea) for Horses
Deficiency
Excess
Diet-Induced Allergy
Carbohydrates
Types and Utilization
Dietary Fiber
Fats

NUTRIENTS

A nutrient is any feed constituent that is necessary for the support of life. Nutrients accomplish this in the following ways:

1. By serving as constituents of the body.
2. By enhancing or being involved in chemical reactions that occur in the body, i.e., body metabolism.
3. By serving as a source of energy.
4. By transporting substances into, throughout, or out of the body.
5. By assisting in the regulation of body temperature, for both heat production and dissipation.
6. By affecting feed palatability and, therefore, consumption.

There are six basic classes of nutrients: (1) water, (2) proteins, (3) carbohydrates, (4) fats, (5) minerals, and (6) vitamins. Some nutrients fill a number of these life-support functions. For example, water and several minerals are needed for all of the functions, except as sources of energy. Proteins, carbohydrates, and fats may all be used for energy but are also constituents of the body. In contrast, vitamins serve only one function: they are necessary for body metabolism.

The major sources, needs, and functions, and the causes and effects, and diagnoses of inadequate and excessive intake of nutrients by the horse are described in Chapters 1, 2, and 3; for those described in this chapter (water, protein, fiber, fats or fatty acids, and nutrients used for energy), they are summarized in Table 1–1.

Nutrients are, of course, present in feeds, which should be thought of simply as nutrient packages—packages that vary in their appearance and palatability, as well as in the quantity of different nutrients they contain, but little in which nutrients they contain. Except for vitamin and mineral supplements, which may contain only certain specific vitamins or minerals, and oils, which contain only fats, all other feeds contain nearly the entire spectrum of different nutrients. Feeds differ therefore not in which nutrients they contain but in the amount of each nutrient they contain. For example, soybean meal, corn, and hay each contain all the nutrients, but soybean meal contains much more protein and less carbohydrate than corn and hay, and hay contains much more fiber than soybean meal and corn. These, as well as other differences in various feeds for the horse, are described in Chapters 4 and 5.

Regardless of the feed, in order for its nutrients to be utilized, they must be released from the feed that has been ingested, broken down sufficiently by digestion, and absorbed into the body by the digestive tract, as shown in Figure 1–1.

WATER

An adequate supply of good-quality, palatable water is essential for horses. Always ensure that adequate, good-quality, palatable water is readily available for all horses at all times. The only exception is that after exercise, the horse should be cooled down before being allowed to drink as much as it wants. Consumption of excess cold water by a horse that is hot from physical exertion may cause colic or founder. However, just before and during prolonged physical activity, the horse should be allowed to drink as often as practical and as much as it wants.

Water Needs

Voluntary water intake by the horse at rest in a moderate or cool environment, eating dry forage, varies from 0.3 to 0.8 gal/100 lbs body weight/day (25 to 70 ml/kg/day). The amount actually required is near the lower end of this range. At rest water requirement in quarts or liters/day is approximately equal to digestible energy requirement in megacalories/day, which for the horse is given in Appendix Tables 4 and 5.

The amount of water needed varies primarily with the amount of water lost from the body, which is altered by the amount, type, and quality of the feed consumed, the ambient temperature and humidity, and the health, physiological state, and physical activity of the horse. The horse, like all animals, consumes more water than needed if palatable water is readily available. The amount of water consumed, however, will decrease to just meet needs if water is poorly accessible or poorly palatable. The amount of water drunk, but not consumed, also decreases with increasing moisture content of the feed. Feed containing 40% or more moisture supplies enough water to meet the idle horse’s needs in a moderate environment. Although hay, grain, and nongrowing forage contain less than 15% moisture, growing forage contains from at least 60% to over 80% moisture. Thus, the horse consuming growing forage does not need to drink any water, although most will if it’s available and palatable. (Figure 1–2).

TABLE 1–1 Water, Energy, Protein, Fiber and Fat: Causes and Effects of Deficiencies and Excesses

 Nutrient Imbalance Causes Effects
Water deficiency Unavailable or unpalatable water, or ↓. thirst due to excess loss of salts in sweat ↓ Performance ↓ Feed intake Dehydration
Energy excess Food intake > needs due to high paiatability and/or little exercise Hyperactivity (“high”) and/or excess body fat
Energy deficiency Inadequate feed: 1) available, 2) caloric density, or 3) eating or utilization ability Weight loss and/or ↓ growth, performance, or lactation
Protein excess Diet protein > needs (See App. Ts. 1 and 4) ↑ urine volume and heat production
Protein deficiency Protein < needs (see App. Ts. 1 & 4). Low protein digestibility or inadequate feed intake Same as energy deficiency. Rough hair coat and may eat feces
Fiber excess Diet crude fiber >34-40%, depending on energy needs “Hay belly” and same as energy deficiency
Fiber deficiency/nonfiber carbohydrate excess Sudden grain excess or grain >50 to 90% of diet Diarrhea, colic, and founder
Fat deficiency Not known to occur in horses In others, dry, dull hair, scaly skin, hair loss and ↓ reproduction
Fat excess Adding >20% oil to diet or >30% oil to grain mix ↓ paiatability, loose, fatty appearing stools. Long-term it results in excess energy

App. = Appendix; T. = Table; ↑ = increase; ↓ = decrease; > = greater than; < = less than.

The amount of water drunk directly correlates with the amount of feed dry matter consumed. Although horses generally drink 1.5 to 2 quarts of water per lb. (3 to 4 L/kg) of hay or grain only about 1 quart/lb (2 L/kg) are normally needed. Donkeys generally drink 1.2 to 2.6 and Shetland Ponies 2.2 to 2.5 L/kg, which may reflect at least the donkey’s desert origin. Water intake decreases with increasing diet digestibility, because increasing diet digestibility decreases the amount of feces and, therefore, the amount of fecal water excreted. Diet digestibility increases, and therefore the amount of water per unit of feed dry matter consumed decreases, with increasing forage digestibility and as the amount of grain in the diet increases.

Water needs and intake also increase with increasing protein and salt intake. Increased protein intake increases nitrogenous waste products excreted in the urine, and this, like increased salt intake, increases urine volume.

Lactating mares have increased water needs to compensate for increased water loss in the milk. Lactation may increase water needs, and also dietary energy needs 1.5 to 1.8 times that required for maintenance. There is also increased water needs during growth and the last trimester of pregnancy. Although a small amount of increased water need for growth is caused by increasing body size and, in pregnancy, placental fluids and the fetus, most of the increased water need during growth, pregnancy, and lactation is due to increased feed intake.

One of the major factors affecting how much water the healthy horse needs is how much water the horse loses through sweat and expired air in order to prevent the body’s overheating from physical activity or the environment. The amount of water needed may increase as much as 3 to 4 times with work at high ambient temperatures. Moderate work alone may increase water needs 1.6 to 1.8 times and hard work 2.2 times that needed at rest. At an ambient temperature of 0°F (−18°C), horses consume 1 qt water/lb (2 L/kg) dry feed eaten, whereas at 100°F (38°C) they will consume four times this amount. An increase in temperature from only 55°F (13°C) to 70°F (21°C) increases the horse’s water requirements by 15 to 20%.

Fig. 1–1. Gastrointestinal tract (GIT) of the horse. The stomach of the 1100 lb. (500 kg.) horse holds 2 to 4 gallons (7.5 to 1 5 liters). Some protein digestion and partial breakdown of the feed occur in the stomach. Liquids pass from the stomach rapidly with 75% gone within 30 minutes after ingestion. Of the feed dry matter ingested, only 25% is gone from the stomach by 30 minutes, and more than 98% by 12 hrs following its ingestion. Although solid particles are partially broken down in the stomach by the acid, and protein by pepsin which it secretes, little digestion occurs in the stomach. In addition, in contrast to most animals, the horse cannot vomit or regurgitate material from the stomach. Most of the feed dry matter ingested passes as particulate matter to the small intestine. The small intestine is 50 to 70 ft (15 to 22 m) long, 3 to 4 inches (7 to 10 cm) in diameter, and holds 10 to 12 gal (40 to 50 L). Much of the fat and protein, and about 50 to 70% of the soluble carbohydrate or nitrogen free extract are digested in the small intestine. These and most of the vitamins and minerals are absorbed from the small intestine. Liquids pass through the small intestine rapidly, and reach the cecum 2 to 8 hours after ingestion. In another 5 hours, most of the liquid that reaches the cecum passes on into the colon. Passage of both liquids and particulate matter through the colon is slow and occurs over a period of about 36 to 48 hours. Nearly all of the crude fiber or cellulose and much (over 50%) of the soluble carbohydrate in feeds passes through the small intestine into the cecum. The cecum is 3 to 4 ft. (0.9 to 1.2 m) long and holds 7 to 8 gal (25 to 30 L). It, like the ascending colon, contains bacteria that digest much of the fiber and about one-half of the soluble carbohydrate (NFE) ingested. After digestion, these nutrients are absorbed from the cecum and colon. Some bacterial protein is also produced, digested and absorbed from the cecum and colon. The large, or ascending, colon is 10 to 12 ft (3 to 3.7 m) long with an average diameter of 8 to 10 inches (20 to 25 cm) and holds 14 to 16 gal (50 to 60 L). It consists of four portions: (1) the right ventral colon, (2) the sternal flexure to the left ventral colon, (3) the pelvic flexure (where obstruction most commonly occurs) to the left dorsal colon, and (4) the diaphragmatic flexure to the right dorsal colon, which connects to the small colon. The small colon is about 10 ft (3 m) long, 3 to 4 in (7.5 to 10 cm) in diameter, and holds about 5 gal (18 to 19 L). When it enters the pelvic inlet, it is called the rectum, which is about 1 ft (0.3 m) long and opens to the exterior at the anus. The large colon, small colon, and rectum make up the large intestine. The empty GIT constitutes 4.2 to 5.2%, the liver 1.1 to 1.4%, and the pancreas 0.9 to 1.0% of the mature horse’s body weight. All decrease with exercise, probably because of blood shunted away from them. All increase when grain is fed, particularly the small intestine. The GIT is smaller and the liver larger in foals, with each constituting 3.5% of body weight at birth and being the same as the adult’s at six months of age. The foal’s GIT is smaller because of an undeveloped large intestine. Intestinal tract length increases rapidly in the fetus, and foal, from mid-gestation to 1 yr of age, and changes little thereafter. Small intestine length increases rapidly during the first month of life, while large intestine length increases most when forage consumption increases.

image

Fig. 1–2(A,B). Waterers that automatically fill when their water level is lowered. They may have a thermostatically controlled heater to prevent the water from freezing during cold weather. Although many automatic waterers’ basins hold only 1 to 2 gal (4 to 8 L), they refill rapidly. If horses always have an automatic waterer readily available, one is generally enough for a number of horses, even during hot weather, because regardless of water needs, horses drink a relatively small amount at one time. With increased water needs, horses drink more frequently, not longer nor with more than generally 1 or 2 drinks each drinking bout. This may not be the case, however, if the horses have access to water for only specific short periods of time. Feed and other debris should be removed from water containers daily, and they should be thoroughly cleaned frequently. They should be placed away from the feed-bunk to minimize their contamination with feed (A). A double waterer (B) may be placed between two stalls or two paddocks.

image

When water is readily available, increased water consumption occurs as a result of increased drinking frequency, not increased drinking duration or the number of drinks taken during a drinking bout. For example, there is a direct correlation between drinking frequency and ambient temperature, with a large increase in frequency at temperatures above 85°F (30°C). When water is readily available, most horses drink once for only about 30 seconds or less every few hours. However, if water is not readily available such as if there is a long distance between preferred grazing areas and water, more and longer drinks may be taken during a drinking bout.

Water Deficiency

Inadequate water intake is quite detrimental. With the exception of inspired oxygen, a deficiency of water produces death more rapidly than a deficiency of any other substance. The first noticeable effect of inadequate water intake is decreased dry feed intake, followed by decreased physical activity and ability. Inadequate water intake is also believed to increase the risk of intestinal impactions and colic. Water deprivation for 24, 48, and 72 hours decreased the normal resting horse’s body weight 4%, 6.8%, and 9%, respectively, when the ambient temperature was 63-81°F (17-27°C). At an ambient daytime maximum temperature of 104°F (40°C), body weight decreased 11 to 13% after 60 hours, and 14 to 16% after 72 hours of water deprivation. Signs of dehydration, such as dry membranes and mouth and sunken eyes, are not evident until at least a 6% loss of body weight has occurred. Less than one-half this amount of dehydration is likely to decrease physical performance. Thus, the horse’s physical performance ability is decreased long before a water deficiency induced dehydration can be detected from the horse’s appearance.

Inadequate water intake occurs when water is poorly palatable or accessible. Palatability is best determined by tasting the water and, if there has been a change in the water or its source, comparing its taste to that to which the horse is accustomed. Poor palatability may be due to poor water quality. Water may be poorly accessible for many reasons, such as if electric heaters with wiring problems cause the animal to be shocked when attempting to drink, or if water is frozen over. Ambient temperature-induced variations in water temperature may not alter water intake. Although this situation has not been studied for horses, cattle drink similar amounts of cold or warm water, although individual cattle or horses may have a preference. Cattle, and therefore possibly horses, will consume sufficient snow or crushed ice to meet their water needs if snow or ice is available but water isn’t. However, in doing so, the total amount of water and feed consumed will be reduced.

Water Quality

The single most reliable indication of water quality is the amount of total dissolved solids (TDS) in the water. The amount of TDS, as given in Table 1–2, provides a useful overall indication of the suitability of a particular water source for livestock use. Water high in TDS may be undesirable or unfit for consumption. This occurrence is most prevalent in arid areas, such as the western non-coastal part of the United States. A TDS of 6,500 ppm (parts per million or mg/L) is considered the upper safe limit in water for horses.

The amount of TDS is the sum of the concentrations of all substances dissolved in water. The term “salinity” as applied to fresh water is often used synonymously with TDS. Another term used to described water quality is total alkalinity, but this is not as good an indication of water quality as is TDS. Total alkalinity is the sum of the concentrations of alkali metals, which are primarily sodium and potassium, but may also include lithium, rubidium, cesium, and francium. The hydroxides of these metals are alkaline; i.e., in water they neutralize acids. The total alkalinity of water is always less than its TDS, or salinity, since TDS and salinity include the sum of the concentrations of all substances dissolved in water, and total alkalinity includes only the sum of the concentrations of alkali metals. Salinity and TDS should not be confused with hardness. Highly saline waters may contain low levels of the minerals responsible for hardness. Water “hardness” indicates the tendency of water to precipitate soap or to form a scale on heated surfaces. Hardness is generally expressed as the sum of calcium and magnesium reported in equivalent amounts of calcium carbonate. Other substances, such as strontium, iron, zinc, and manganese, also contribute to hardness.

TABLE 1–2 A Guide to the Suitability of Water for Livestock

TDS (ppm)a Suitability and Effect
1000-3000 Satisfactory for all livestock and poultry. May cause mild and temporary diarrhea in livestock not accustomed to it, but should not affect their health or performance.
3000-5000 Should be satisfactory for livestock, although it might cause temporary diarrhea, or be refused at first by animals not accustomed to it.
5000-7000 Can be used with reasonable safety for livestock. May be advisable to avoid water approaching the higher level of ppm for pregnant or lactating animals.
7000-1 0,000 Unfit for poultry and swine. Considerable risk may exist in using this water for pregnant, lactating, or young animals, or for any animals subjected to heavy heat stress or water loss. In general, the use of this water should be avoided, although animals other than those listed here may subsist on it for long periods.
Over 10,000 Not recommended for use by any animal under any condition

a Total dissolved solids, total soluble salts, or salinity in the water in ppm or mg/L

Sodium, potassium, calcium, magnesium, iron, chloride, and sulfate in water are not toxic, but high concentrations decrease water palatability. In contrast, a number of other substances, which may be present in water, are quite toxic if sufficiently high concentrations are present. Toxic concentrations of water contaminants, excluding pesticides and herbicides, most commonly occur as a result of stagnant or runoff water that contains disease-producing organisms, or from industrial wastes. A list of the recommended upper limits for some potentially toxic substances in drinking water for horses, and those not toxic but which, if present at concentrations above those given, reduce water palatability, is given in Table 1–3. Some potentially toxic substances do not reduce water palatability and, therefore, water intake. Thus, they are potentially even more harmful than those that do decrease palatability. In addition to these contaminants, drinking water containing some bacteria and algae may be harmful.

Some species of blue-green algae, which grow on pond and lake water, may result in poisoning; therefore, water with heavy algae growth should be avoided. Heavy algae growth occurs most commonly during summer and fall in shallow, still water rich in organic nutrients. These nutrients may be increased, and thus algae growth promoted, by runoff of nitrogen or phosphate from slurry lagoons, or of fertilizers applied to fields. Steady prevailing winds may concentrate the algae at one end of the pond or lake, increasing the risk of poisoning. The algae may be visible on the water surface or mixed with the water.

Blue-green algae poisoning in domestic livestock may cause sudden death or else photosensitization, tremors, weakness, bloody diarrhea, and convulsions. Clumps of algae may be found in the gastrointestinal contents of animals that die suddenly. Copper sulfate added to pond water up to a concentration of 1 ppm (1 mg/L) has been used successfully to kill algae blooms, but will probably be harmful to other types of aquatic life.

Water high in bacteria is usually also high in nitrates as a result of surface contamination from manure and barnyard runoff. However, high nitrate water levels may come from other nitrate sources, such as crop fertilizers, and not be high in bacteria. Nitrates may build up in well water by leaching down through the soil. Water nitrate levels may fluctuate widely; they are generally highest following wet periods, and lowest during dry periods of the year. Since nitrates dissolve in water, they cannot be filtered out; however, commercially available anion exchange units remove both nitrates and sulfates. Nitrate toxicosis, however, as described in Chapter 19, is rare in horses, if it occurs at all, and in livestock is most often associated with high nitrate levels in forage, not water. Water sulfate concentrations exceeding 1000 ppm may cause diarrhea, although animals develop a tolerance to a constantly high level of sulfates and can tolerate two to three times this concentration after a period of time. Water with low levels of sulfates, however, may have an odor and reduced palatability.

In most areas and situations, bacteria in water pose a greater threat than the contaminants previously discussed and listed in Table 1–3. Most infectious diseases can be transmitted from contaminated water to animals. If water nitrate or phosphate concentrations are low, the water probably does not contain excessive bacteria. However, if either is high, bacterial levels may be elevated and should be checked. The accepted criterium for the sanitary quality of water is the absence of coliform bacteria. Although all coliform bacteria are not disease producing, many are, and their presence indicates that other infectious bacteria and viruses may be in the water. The U.S. Public Health Service considers water containing coliform bacteria (M.P.N.) of 9 or more coliforms per 100 ml unsafe for human consumption. In some countries, levels of 50 coliforms/100 ml are acceptable. What amount is safe for horses isn’t known but, of course, also depends on which organisms are present.

TABLE 1–3 Recommended Upper Safe Level (USL) of Water Contaminants

Contaminate USLa
Arsenic 0.2
Cadmium 0.05
Calcium 500b
Chloride 3000b
Chromium 1
Cobalt 1
Copper 0.5b
Cyanide 0.01
Fluoride 2c
Hardness 200
Hydrogen Sulfide 0.1
Iron 0.3b
Lead 0.1
Magnesium 125b
Manganese 0.05
Mercury 0.01
Nickel 1.0
Nitrate (see Ch. 19) 400 ± 10d
Nitrate nitrogen 100
Nitrite nitrogen 10
Potassium 1400b
Selenium 0.01e
Silver 0.05
Sodium 2500b
Sulfate 2500f
TDS (see Table 1–2) 6500
Vanadium 0.1
Zinc 25g

aAll values given are in parts of contaminate per million parts of water (ppm or mg/L). For conversion to other units, see Appendix Table 9.

b These contaminates are not toxic but at concentrations above the amount given may decrease water palatability. In contrast, many of the other contaminates listed may be toxic if water containing concentrations above those given here is the only water consumed.

c A higher concentration may be safe for horses, as 2.5 ppm results in mottled enamel during teeth development in calves but no observable effects occur in mature cattle at concentrations of less than 8 ppm, and horses are reported to tolerate fluoride intakes two to three times greater than cattle. A concentration of 4 ppm is probably marginally safe for horses, but water with more than 8 ppm should be avoided.

d High nitrate concentrations in water occur most commonly as a result of fecal contamination.

e Although chronic selenium toxicosis has been reported as a result of consumption of water containing 0.0005 to 0.002 ppm selenium, concentrations below 0.01 ppm are not generally considered harmful.

f Or 833 ppm sulfur. Although sulfate concentrations above 300 to 400 ppm can be tasted, and above 750 ppm can have a laxative effect in people, a concentration below 2500 ppm has no effect on growing or reproducing cattle or swine.55 The highest no-effect concentration in horses isn’t known but is probably similar to that for cattle and swine.

g High zinc concentrations may occur where galvanized pipes are connected to copper. This results in electrolysis, releasing zinc from the galvanized pipes into the water.

Salmonella species is generally the bacterial contaminant in water most likely to cause disease in farm animals. Giardia is the most common cause of water-related illness in people, partly because it survives chlorination. Although giardiasis is rare in farm animals, it can cause diarrhea in young animals. The most common method of destroying bacteria in a water supply is chlorination, although iodine, ozone, exposure to ultraviolet rays or ultrasonics, or filters may be used. Objectionable chlorine taste and odor can be removed from water by an activated carbon filter.

In summary, flowing surface water is most likely to have bacterial contamination, pond or lake water is most likely to contain blue-green algae, and well water, particularly in arid areas, is most likely to have high mineral concentrations. Coliform counts and measures of total dissolved solids are the main indications of water quality.

DIETARY ENERGY

Energy Sources and Use

As described by the first law of thermodynamics, energy can be changed from one form to another but can be neither created nor destroyed. The source of all energy for all living things is sunlight. This energy is captured by plants, which use it via photosynthesis to change carbon dioxide, present in the air, and water to oxygen and the carbon compounds that make up the plant. These carbon, or organic, compounds in plants are carbohydrates, fats, and, along with nitrogen, protein. Nitrogen, like the small amount of minerals needed by plants, is taken up from the soil but is originally from the air. Carbohydrates, fats, and proteins are stored sources of energy.

Animals eat plants, or tissues of others that have eaten plants. The stored sources of energy, carbohydrates, fats, and proteins in the plants are digested, absorbed, and transported to the animal’s body cells. Some are used to make up the structural components of the cell and thus the animal; but, if needed in the future, along with oxygen from the air, they can be converted by chemical reaction to carbon dioxide and water, in the process producing energy. Animals use the energy to produce heat and adenosine triphosphate, or ATP, which cells then use to function. Thus, plants and animals have a mutually sustaining relationship in which plants produce carbon compounds (C-cpds)—organic matter and oxygen (O2)—that support animals, and animals produce the carbon dioxide (CO2) and water (H2O) that support plants Thus, the horse, like us, plants, and all living things on earth, are products of the air and the soil, and function solely as the result of solar energy.

Energy has no measurable dimension or mass, but it can be converted to heat, which can be measured. Oxidation. or burning, converts stored energy (carbohydrates, fats, and proteins) to heat, carbon dioxide, water, and a nitrogen compound. These are returned to the air and soil, where they originated, available to begin the cycle again. When a substance is completely oxidized, the heat produced, called the heat of combustion, is the total or gross amount of energy stored in, and thus available from, that substance. However, as shown in Figure 1–3, the animal cannot use all of the gross energy present in a feed. Some of the stored energy in a feed is not digested and is lost in the feces. Of the remainder, called digestible energy, some is lost in the urine as urea and in the gastrointestinal tract as gases (primarily methane), leaving metabolizable energy, of which some is used in metabolizing feed. What remains is the net energy available for maintenance, growth or fattening, milk production, and physical activity. Most of the total dietary energy needs are for maintenance. Even during heavy lactation or physical activity over 50% of dietary energy is needed for maintenance, and for young horses, 60 to 95% of dietary energy needs are for maintenance, leaving 5 to 40% for their growth.

Fig. 1–3. Dietary Energy Partition.

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The amount of heat produced by oxidation (burning) that raises the temperature of one gram of water 1°C is defined as one calorie. This is also known as the small, gram, or standard calorie. However, it is not used in nutrition for animals or people. The calorie used in nutrition is the amount of heat required to raise one kilogram of water 1°C. It is called the large calorie, Calorie, or kilocalorie (kcal) since it is equal to 1000 small calories. In nutrition, the word calorie always refers to kilocalorie, even if it is not capitalized and neither the kilo- or k-prefix is used. For large animals, such as horses, megacalorie (Mcal), therm, or total digestible nutrients (TDN) are usually used. One megacalorie equals one therm, and both equal 1000 kilocalories. Occasionally, primarily in England, instead of calorie, the joule, or, in the physical sciences, the British Thermal Unit (BTU) is used. One megacalorie equals 4.1855 megajoules and 3968 BTU. TDN is a measure of digestible energy expressed in units of weight or percent, with 1 lb TDN equal to about 2.0 Mcal DE (1 kg TDN = 4.4 Mcal). TDN is the sum of digestible carbohydrates, plus digestible protein, plus digestible fats times 2.25, because fats provide about 2.25 times more energy than an equal weight of carbohydrates or proteins. Starch equivalent, or SE, is occasionally used as an energy term by comparing the energy provided by a feed to that provided by starch, which is assigned a value of 100%. Although occasionally used for ruminants, SE is unsuitable for horses because of their different digestive process.

When calculating energy intake, or the amount of feed needed to provide a certain amount of energy, such as that necessary to meet the animal’s energy requirements, any of the various energy terms may be used. Of course, the same units must be used for both the energy content of the feed and the animal’s energy needs. Net energy is the most accurate, followed by metabolizable energy (Fig. 1–3). However, they are the most difficult to determine and, as a result, are not routinely available for most horse feeds. Digestible energy values (or TDN) are generally available for most horse feeds and therefore are the most commonly used energy terms. Energy available from forages is usually 5 to 15% higher for cattle than for horses because of ruminants’ more efficient utilization of fiber. Therefore, if the energy content of forages for cattle, or other ruminants, is used to determine the amount of these feeds needed by horses, the amount determined generally will be erroneously low.

Energy Needs

Numerous factors can influence the energy requirements of the horse. These include environmental conditions, the horse’s functions and activity (including intensity and duration of work, weight and ability of the rider, and conditions of the traveling surface), and its physical condition and degree of fatigue. Even when all of these factors are identical, individual horses vary in their energy needs. The average amount of energy needed, as given in Appendix Tables 1, 4, and 5, should therefore be considered only a general guideline—an amount that will be relatively close for a group of horses but either inadequate or excessive for some individuals.

The amount of digestible energy (DE) needed for maintenance (i.e., for no weight change by the mature, idle nonreproducing horse at moderate environmental conditions) by the average horse weighing 1320 lbs (600 kg) or less can be calculated from the following equation.

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However, energy needs are lower per unit of body size in horses weighing over 1320 lbs (600 kg) and can be calculated from the following equation.

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Estimates of energy requirements for physical activity or work depend primarily on the total weight carried (horse, intestinal fill, rider, and tack) times the distance moved, but increase with decreasing ability of the rider and physical condition of the horse, difficulty of the terrain and surface covered, and other factors. For ponies and light horses, the Mcal DE/day for light, medium, and intense work has been estimated to be respectively 1.25, 1.50, and 2.0 times that needed for maintenance (Appendix Table 4); with light work being activities such as Western and English pleasure, bridle path, hack, and equitation; medium work being ranch work, roping, cutting, barrel racing, and jumping; and intense work being race training, endurance racing, and polo. Digestible energy needs greater than those required at rest have also been estimated in Mcal/hr/100 kg (220 lbs) total weight carried to be 0.17 for a slow walk; 0.25 for a fast walk; 0.6 for a slow trot; 1.0 for a medium trot or slow lope; 1.3 for a fast trot; 2.0 for cantering, galloping or jumping; and 3.9 for a fast gallop (Appendix Table 5).